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       Faunal 
        Remains 
      
      by Robert J. Muir and Jonathan C. Driver 
      
      Introduction
      1 
        In this chapter, we report on the analysis of faunal remains collected 
        during excavations at Yellow Jacket Pueblo (Site 5MT5) by the Crow Canyon 
        Archaeological Center. The assemblage includes all nonhuman bones and 
        teeth, antler, eggshell, and ossified cartilage found at the site. Both 
        modified and unmodified materials were analyzed and are reported on here. 
      2 
        The chapter begins with a brief discussion of the methods used to identify, 
        catalog, and quantify the faunal remains, followed by a description of 
        the animal taxa identified in the assemblage. Much of the chapter is given 
        to consideration of the taphonomic processes that might have influenced 
        the composition of the assemblagethat is, the various natural and 
        cultural processes related to deposition and preservation that likely 
        affected the kinds of remains that were recovered, as well as the condition 
        in which they were found. This discussion includes an assessment of the 
        natural and cultural modifications evident on the specimens and a reporting 
        of the skeletal representation of the most common taxa. Following the 
        discussion of taphonomy is an intrasite analysis of the spatial and chronological 
        distribution of the faunal remains, by architectural block. Finally, we 
        compare the faunal assemblage from Yellow Jacket Pueblo with the assemblages 
        from other Pueblo III sites in the region in an attempt to discern similarities 
        and differences in intrasite spatial patterning that might reflect economic 
        and/or social variation among communities. 
      Methods: Identification, Recording, and Quantification of Faunal Remains
      3 
        All faunal remains collected during the excavations were analyzed by the 
        authors and cataloged using a standardized identification and recording 
        system developed by Jon Driver for Crow Canyon (Driver 
        1992*2). The following information was recorded for each specimen: 
        taxon, element, part of element, side, state of epiphyseal fusion, type 
        of breakage, modifications (cut marks, grinding, burning, weathering, 
        gnawing, etc.), length of fragment, and cortical thickness. Identifications 
        were made using comparative collections at Simon Fraser University, the 
        University of Puget Sound Museum of Zoology, and the Burke Memorial Museum 
        (Washington). We also used several osteological keys (including Gilbert 
        et al. 1981*1; Lawrence 
        1951*1; Olsen 1964*1, 
        1968*2; Schmid 
        1972*1) to assist us in sorting and making preliminary identifications. 
        Those interested in a more detailed description of identification and 
        cataloging procedures should consult additional sources (Driver 
        1992*1, 1992*2; Driver 
        et al. 1999*1).  
      4 
        Before identifying and cataloging the remains, we made a considerable 
        effort to reconstruct elements that had been broken during or after excavation. 
        We made no attempt to reconstruct elements that displayed old breaksthat 
        is, breaks that occurred before excavation. Fragments that were obviously 
        parts of the same bone were noted in the catalog. By preserving fragmentation 
        caused by cultural or natural taphonomic processes, we were able to view 
        the assemblage as it would have existed just before excavation. In the 
        subsequent analysis, each fragment or reconstructed element was counted 
        as a distinct specimen. 
      5 
        During analysis, a specimen was considered "identifiable" only if the 
        skeletal element could be positively determined. General element categories 
        such as "long bone" or "axial" were not used. All specimens that could 
        not be identified to a specific element were thus classified taxonomically 
        as "unidentifiable." This requirement ensures that analyses are not unduly 
        biased by the intuition and guesswork of individual analysts (Driver 
        1992*1). 
      6 
        All identifiable specimens were assigned to the most specific taxonomic 
        category possible, given the limitations of the available reference collections 
        and observable morphological variation. Bones were assigned to a species 
        or genus only when all other possibilities had been examined and ruled 
        out on the basis of morphology and size. Species-level identifications 
        were made only by direct comparison with modern skeletons. Many specimens 
        were assigned to more general taxonomic categories as defined by Driver 
        (1992*2). 
      7 
        Frequency data for the faunal remains from Yellow Jacket Pueblo are provided 
        as "number of identified specimens," or NISP counts (Grayson 
        1979*1). NISP counts represent the total number of specimens that 
        can be positively identified as belonging to a particular taxon. This 
        method has a number of potential problems (for a thorough discussion, 
        see Grayson [1979*1]). In 
        particular, NISP data will overrepresent taxa with (1) greater numbers 
        of elements (Klein and Cruz-Uribe 
        1984*1; Payne 1972*1); 
        (2) greater degrees of fragmentation (Grayson 
        1973*1, 1979*1; Thomas 
        1969*1; Watson 1979*1); 
        or (3) higher rates of recovery (Thomas 
        1969*1; Watson 1972*1). 
        Furthermore, NISP counts will produce artificially inflated sample sizes 
        (Watson 1979*1). NISP data 
        have been used here to allow direct comparison to faunal data produced 
        by other researchers, but they do not provide a particularly precise estimate 
        of taxonomic abundance. 
      Identified Taxa
      8 
        Crow Canyon's test excavations resulted in the collection of 9,132 bone, 
        tooth, antler, and ossified cartilage specimens from the site. Of these, 
        3,939 could be identified to element and thus assigned to a specific taxonomic 
        category (Table 1). Approximately 
        200 bird eggshell fragments were collected from the site but were not 
        included in the analyses and discussion presented in this chapter. 
      9 
        Mammal remains dominate the identifiable assemblage, accounting for 69.0 
        percent of the specimens. Birds represent approximately one third (30.8 
        percent) of the identified remains, and trace amounts of fish and reptile 
        remains make up the balance of the assemblage. The identified specimens 
        represent a minimum of 40 mutually discrete taxonomic groups, including 
        at least 28 mammal and nine bird taxa (as well as one fish and two reptile 
        taxa). Additional taxonomic categories may be represented in the remains, 
        as a considerable number of specimens have been assigned to general categories 
        such as "medium carnivore" and "large bird." The majority of these remains 
        probably belong to taxa already identified within the assemblage. For 
        example, most specimens identified as "large bird" are probably the remains 
        of Meleagris gallopavo (turkey) or of one of the other large 
        bird species already identified within the assemblage. Similarly, remains 
        identified as "medium artiodactyl" are undoubtedly of Odocoileus 
        sp. (deer), Antilocapra americana (pronghorn antelope), or 
        Ovis canadensis (bighorn sheep). On the contrary, the wide variety 
        of Muridae (deer mice, voles, etc.) is difficult to separate osteologically, 
        and it is possible that species in addition to those named are represented 
        by these remains. Similarly, remains identified as "Sciuridae" (squirrels) 
        may represent a number of species not listed. 
      Mammals
      10 
        The mammal remains from Yellow Jacket Pueblo include a wide variety of 
        taxa, although many are represented by only a few specimens (Table 
        2). The lagomorphs are most common, representing more than 54 percent 
        of the mammalian subassemblage. Cottontails (Sylvilagus sp.) are 
        extremely abundant, whereas jackrabbits (Lepus sp.) are represented 
        in much smaller numbers. Two species of cottontail may be represented: 
        Sylvilagus audubonii and Sylvilagus nutallii. No attempt 
        was made to distinguish between these species. The Lepus remains 
        were not assigned to a species, although snowshoe hare (Lepus americanus) 
        can be ruled out on the basis of size. No pikas (Ochotona sp.) 
        were positively identified among the remains. 
      11 
        A considerable quantity of rodent remains was recovered from the site, 
        representing approximately 24 percent of the mammalian specimens. Most 
        of the elements that were identified to genus and species were mandibles, 
        crania, teeth, innominates, and major long bones. Other rodent elements 
        have been identified only to the family level. Small rodents, including 
        deer mice and voles (Muridae), woodrats (Neotoma sp.), and pocket 
        gophers (Geomyidae), are most numerous. These animals are probably underrepresented, 
        given the potential for their very small bones to be lost or overlooked 
        during excavation. The larger rodents include rock squirrel (Spermophilus 
        variegatus), prairie dog (Cynomys sp.), porcupine (Erethizon 
        dorsatum), and beaver (Castor canadensis). Single specimens 
        representing chipmunk (Eutamias sp.) and Abert's squirrel (Sciurus 
        aberti) round out the rodent remains. 
      12 
        The order Carnivora is represented by at least five taxa and makes up 
        2.4 percent of the mammalian remains. Canids (fox, coyote, dog, and wolf) 
        are by far the most common family of carnivore represented; only one species, 
        domestic dog (C. familiaris), was positively identified. Other 
        carnivores are represented in smaller quantities, including marten (Martes 
        americana), long-tailed weasel (Mustela frenata), badger (Taxidea 
        taxus), and lynx (Lynx sp.). The lynx remains may be of bobcat 
        (L. rufus) or Canada lynx (L. canadensis), although the 
        latter is less probable, given this species' preference for heavily forested 
        environments and northern latitudes (Wooding 
        1982*1:130132). 
      13 
        Artiodactyl remains account for less than 9 percent of the mammalian assemblage. 
        At least five species are represented: deer (Odocoileus sp.), 
        pronghorn antelope (Antilocapra americana), bighorn 
        sheep (Ovis canadensis), elk (Cervus elaphus), and domestic 
        cattle (Bos taurus). Most of the artiodactyl remains were identified 
        simply as "medium artiodactyl." Among those remains that could be identified 
        more precisely, deer (Odocoileus sp.)representing either 
        mule deer (Odocoileus hemionus) or white-tailed deer (Odocoileus 
        virginianus)are most abundant. 
      14 
        Finally, the orders Perissodactyla and Insectivora are each represented 
        by a single specimen: a horse (Equus caballus) radius and a shrew 
        (Soricidae) mandible, respectively. 
      Birds
      15 
        Two taxaMeleagris gallopavo (turkey) and "large bird"dominate 
        the bird (Aves) subassemblage; together these make up more than 90 percent 
        of the bird remains (Table 3). 
        Many of the "large bird" bones are probably turkey. Birds such as owls, 
        ravens, falcons, and hawks may also be represented by the "large bird" 
        category, though probably only in very small proportions. In addition 
        to turkey, small quantities of other Galliformes (including quail and 
        sage grouse) were identified. 
      16 
        The order Columbiformes (pigeons and doves) is represented by two specimens, 
        or 0.16 percent of the bird subassemblage. Identifications to the family 
        or genus level were not possible for these remains. 
      17 
        Birds of prey, including members of the orders Falconiformes and Strigiformes, 
        are also present in the Yellow Jacket assemblage (1 percent). Among these, 
        the remains of hawk (Buteo sp.), falcon (Falco sp.), 
        and short- or long-eared owl (Asio sp.) were identified. 
      18 
        Passerine birds make up less than 1 percent of the bird subassemblage. 
        A single raven (Corvus sp.) specimen is conspicuous and easily 
        identified because of its relatively large size. The common raven (C. 
        corax), which is native to the area, is likely the species represented, 
        but the smaller Chihuahuan raven (C. cryptoleucus) could not 
        be excluded. The other Passeriformes remains are clearly those of smaller 
        species. 
      Reptiles and Fish
      19 
        NISP data for the nine reptile and fish specimens recovered from Yellow 
        Jacket Pueblo are presented in Table 
        4. The reptile subassemblage contains both lizard and snake remains. 
        Four fragmentary fish bones were also identified. Little effort was made 
        to specifically identify any of these specimens because the comparative 
        collections used did not include a complete range of species. 
      Taphonomy
      20 
        Taphonomy is the study of the natural and cultural processes that affect 
        the deposition and preservation of organic materials. Cultural behavior 
        associated with artifact manufacture, food preparation, and refuse disposal 
        will influence the distribution and composition of archaeological assemblages. 
        Once deposited on the ground, objects may be moved by natural mechanical 
        forces such as wind, water, or burrowing animals. Other natural agents 
        such as moisture, sunlight, and bacteria may erode or destroy materials. 
        Soil chemistry also affects preservation. Bones and other organic materials 
        are especially vulnerable to alteration, damage, or destruction as the 
        result of such factors. In this section, we examine evidence of taphonomic 
        processes that have affected the composition and condition of the Yellow 
        Jacket Pueblo faunal assemblage. 
      Weathering of Remains
      21 
       Animal bones can be damaged or destroyed 
        by exposure to moisture, wind, and sun; the effects of such weathering 
        may have a substantial impact on the composition of a faunal assemblage. 
        The primary variable affecting the degree of bone weathering is the rate 
        of burial: bones that become buried more quickly will be less affected 
        by the natural elements. Bones will also weather differentially depending 
        on various natural attributes of the bones, including density, size, and 
        grease content. In addition, cultural practices such as processing, cooking, 
        and disposal may influence a bone's susceptibility to weathering. For 
        any of these reasons, bones of certain species may be more likely to suffer 
        from weathering and, thus, less likely to be recovered and/or identified 
        at an archaeological site. 
      22 
        In the Yellow Jacket Pueblo faunal assemblage, it is evident that some 
        taxa have been more severely affected by destructive weathering processes 
        than others (Table 5). In particular, 
        specimens from many of the medium and large mammalian taxa display evidence 
        of damage due to weathering. Weathering is also apparent on small percentages 
        of the "large bird," Meleagris gallopavo, and lagomorph remains. 
        Weathering is not apparent among the vast majority of small mammal remains, 
        and no medium or small bird remains show evidence of weathering. It is 
        perhaps most interesting that no small rodent remains appear weathered, 
        despite their relative abundance in the assemblage. The variability in 
        degree of weathering seen here may be due to both cultural and natural 
        processes. Whatever the cause, it does appear that the remains of some 
        taxa, particularly the medium and large mammals, are more likely to have 
        suffered from weathering and therefore may be underrepresented in the 
        assemblage. 
      Cultural vs. Natural Bone Accumulations
      23 
        In general, the introduction of animal remains into archaeological sites 
        will occur as the result of one of four processes: (1) death of domesticated 
        animals raised on the site by humans; (2) killing of trapped or hunted 
        animals transported to the site by humans; (3) transport of dead animals 
        to the site by nonhuman predators or scavengers; or (4) natural death 
        of wild animals that lived and died on the site. Whereas the first two 
        processes relate directly to the human occupation of the site, the latter 
        two may be independent of human occupation and thus unrelated to cultural 
        activities; they may occur before, during, or after occupation of the 
        site. 
      24 
        Identification of cultural modification of medium and large mammal remains 
        can usually be made with confidence using discrete criteria such as breakage 
        patterns and the presence of cut marks; this assessment is more difficult 
        for the remains of smaller animals, which can be procured, processed, 
        cooked, and eaten with little modification to skeletal elements. It is 
        therefore impossible to assess how each individual specimen came to be 
        part of the assemblage. By considering the artificially and naturally 
        produced modifications observed on these remains, we hope to determine 
        the most probable taphonomic origin for the specimens of each taxon. 
      Culturally Modified Bone
      25 
        Butchery and processing of animals can produce diagnostic modifications, 
        including cut marks, saw marks, or cut edges. Use of animal bones as tools 
        may also produce modifications such as evidence of grinding and polishing. 
        Remains that display such evidence are obviously indicative of human procurement 
        of the animals that they represent, although such markings are not likely 
        to occur on all, or even a majority, of culturally introduced specimens. 
        Evidence of burning, such as carbonization or calcinization, may be common 
        on culturally utilized taxa, but naturally deposited bones may, of course, 
        become charred as the result of wildfires or accidental house fires (Grayson 
        1988*1; Lyman 1988*1). 
        Spiral fractures are frequently considered indicative of human activity, 
        although these can also be produced by several natural agents, including 
        carnivores (Binford 1981*2), 
        natural traumatic injury (Lyman 
        1984*1), or trampling (Haynes 
        1983*1). 
      26 
        Based on evidence of cultural modification observed in the Yellow Jacket 
        Pueblo faunal assemblage (Table 
        6), a cultural origin for several specific taxa can be inferred with 
        considerable confidence. These include Lepus sp., Sylvilagus 
        sp., Castor canadensis, Canidae, Odocoileus sp., Buteo 
        sp., and Meleagris gallopavo. All display direct cultural modification 
        in the form of grinding, polishing, and cut marks. For some taxa, such 
        modification is extremely common. This is particularly true of the Odocoileus 
        sp. remains, of which 14.3 percent have been made into tools 
        or other artifacts. Numerous specimens in each of these taxa also display 
        burning or spiral fractures. The origins of the remaining taxa are uncertain, 
        as these groups display no definitive evidence of cultural modification. 
      Naturally Modified Bone
      27 
        Bones from animals that were killed and/or consumed by predators and scavengers 
        may display several distinctive modifications. Most obvious are tooth 
        marks left by mammalian carnivores (Binford 
        1981*2), although the bones of very small animals may be totally consumed 
        by predators, such as owls, without extensive modification (Andrews 
        1990*1; Dodson and Wexlar 
        1979*1; Kusmer 1986*1). 
        Bones may also display spiral fractures (Binford 
        1981*2) or be extensively macerated (Korth 
        1979*1) as the result of carnivore predation. The ends of long bones 
        are often gnawed by carnivores and the diaphyses split to obtain marrow. 
        Licking of the ends of long bones by carnivores and other animals may 
        result in smooth, polished surfaces that closely resemble cultural modification 
        (Binford 1981*2; Haynes 
        1980*1). In such cases, crushed bone and tooth marks may be evident 
        elsewhere on the specimen. Acid etching of cortical bone may be evident 
        on bone fragments that have passed through a carnivore's digestive tract 
        (Andrews 1990*1:30). The 
        identification of specimens introduced to the site by carnivores is complicated 
        by the fact that the ancestral Pueblo are known to have kept domestic 
        dogs. Dogs are likely to have had access to discarded animal remains and 
        could have easily produced modifications indistinguishable from those 
        made by wild carnivores. 
      28 
        Table 7 presents the frequencies 
        of carnivore modification to faunal remains from Yellow Jacket Pueblo. 
        Carnivore modification is evident in nine taxonomic groups. It is significant 
        that carnivore modification is relatively common among several taxa (medium 
        artiodactyls, Odocoileus sp., and Meleagris gallopavo) that 
        are almost certainly of cultural origin at the site. This may indicate 
        the presence of domestic dogs. Unfortunately, these data do little to 
        clarify the origin of the other taxa. 
      29 
        Unmodified, predominantly complete bones should result from the natural 
        deaths of animals living on a site (Driver 
        1985*1:18). Burrowing animals may die in their burrows and should 
        display little bone damage from weathering or trampling. Animals that 
        are killed by humans and then processed for food or raw materials are 
        more likely to be represented by fractured specimens, particularly long 
        bones, which are commonly broken to extract marrow. Table 
        8 presents a comparison of complete and fragmented long bones of animals 
        that might have naturally inhabited Yellow Jacket Pueblo and died there 
        in burrows or dens. To mitigate biases due to element size, recovery rates, 
        and identifiability, only major long bones are considered here. Specifically, 
        the humerus, femur, and tibia have been selected for consideration because 
        of their relatively large size and taxonomically diagnostic characteristics. 
        However, it should be noted that differences in the sizes of the animals 
        in these taxa may introduce some biases into the comparisons presented. 
        For example, due to the extremely small size of the long bones of Muridae 
        (deer mice, voles, etc.), only complete specimens are likely to be recovered. 
      30 
        The data presented in Table 8 
        indicate that the Lepus and Sylvilagus long-bone remains 
        are predominantly fragmented specimens, whereas complete elements are 
        more common among the other taxa, particularly the small rodents. As mentioned 
        above, the frequencies may reflect, in part, the influence of animal size 
        on recovery rate, particularly for the smaller rodents; however, the differences 
        between the similarly proportioned Sciuridae and Sylvilagus remains 
        are unlikely to have resulted from factors associated with recovery rates 
        or identifiability. Thirty-three percent of the Sciuridae long bones are 
        complete, whereas less than 5 percent of the lagomorph long bones are 
        complete. The fragmentation data support the argument that the taphonomic 
        histories of the lagomorphs and rodents are considerably different. Specifically, 
        the abundance of complete rodent long bones is consistent with the suggestion 
        that these animals died naturally in their burrows. Conversely, the high 
        frequency of fragmentation among the lagomorph long bones lends further 
        support to the conclusion that these specimens are primarily a product 
        of cultural activity (as suggested by the evidence of cultural modification 
        presented above). Similar patterns have been documented at several other 
        sites in the central Mesa Verde region (Driver 
        et al. 1999*1; Muir 1999*2). 
      Modern Bone
      31 
        Like most archaeological sites, Yellow Jacket Pueblo has been subject 
        to postdepositional disturbance by a variety of agents. Rodent burrowing 
        and other animal activities have undoubtedly been ongoing since establishment 
        of the site. It is also well known that Yellow Jacket Pueblo has been 
        heavily disturbed by artifact collectors, and it is possible that these 
        activities have resulted not only in mixing of deposits and removal of 
        artifacts, but also in introduction of materials to the site assemblage. 
        The remains of two taxa can be confidently assessed as intrusive to the 
        cultural deposits at Yellow Jacket Pueblo: cattle (Bos taurus) 
        and horse (Equus caballus). Neither of these species were present 
        in North America during the occupation of the site. The presence of these 
        specimens confirms that some mixing of modern and prehistoric deposits 
        has occurred. 
      Taphonomic Origins of the Faunal Remains
      32 
        On the basis of the analyses and discussion presented above, we can draw 
        some conclusions regarding the taphonomic origin of the faunal remains 
        at Yellow Jacket Pueblo. Several taxa can be confidently assessed as being, 
        at least in part, the result of human activities. These include all those 
        taxa that display definite indications of cultural modification (that 
        is, cut marks, polishing, or grinding), specifically, Meleagris gallopavo, 
        Odocoileus sp., Canidae, Lepus, Sylvilagus, Castor 
        canadensis, and Buteo sp. It is also possible that a significant 
        number of specimens representing these taxa were introduced to the assemblage 
        naturally. This is most likely for those taxa, such as Sylvilagus, 
        that display only rare indications of cultural modification. 
      33 
        A second group of taxa, consisting of the small- and medium-size rodents, 
        displays characteristics that suggest that their presence is primarily 
        the result of the natural occurrence of these animals at the site. It 
        should be noted that numerous rodent species are mentioned ethnographically 
        as having been trapped and eaten by Pueblo peoples (Gnabasik 
        1981*1). However, no evidence of butchering was observed among the 
        many rodent remains recovered from Yellow Jacket Pueblo. Instead, the 
        rodent remains primarily include complete, unmodified specimens, which 
        is consistent with the interpretation that they represent animals that 
        died naturally in their burrows. The sciurids (squirrels) may be an exception. 
        A higher incidence of long-bone fragmentation than is typical for this 
        group and evidence of spiral fractures and burning suggest that the presence 
        of some of the sciurid remains may be the result of cultural activities. 
      34 
        The origins of the remains of the other taxa are uncertain. Most of these 
        taxa are represented by relatively few specimens, and therefore we have 
        little evidence of their collective taphonomic history. 
      Element Frequencies
      35 
        It is generally expected that most animals will be represented archaeologically 
        by more-or-less complete skeletons, but in some instances cultural and/or 
        natural processes may influence the relative frequencies of particular 
        skeletal regions, elements, or element parts. For example, large game 
        may be represented at a habitation site by only those elements carried 
        there by hunters, or alternatively, at a kill site by only those elements 
        that are left behind (Perkins 
        and Daly 1968*1; White 1953*1). 
        Smaller animals are less likely to be affected by such differential transport 
        of parts, but they may become disarticulated and distributed throughout 
        a site as a result of butchering and processing. Consistent and repeated 
        cultural practices may cause specific portions of some taxa to be selectively 
        preserved, destroyed, or removed from the archaeological assemblage. 
      36 
        Natural agents can have similar effects on frequencies of skeletal parts. 
        Carnivores may selectively remove or destroy specific elements of some 
        species, thus creating assemblages that contain incongruent element compositions 
        (Binford 1981*2:214216; 
        Marean et al. 1992*1). Rodents 
        may collect elements of a particular size range or density, resulting 
        in removal of these elements from a site or their preservation within 
        a burrow (Hoffman and Hays 1987*1). 
        Natural mechanical dispersal mechanisms such as colluvial and fluvial 
        forces may also selectively affect skeletal part frequencies. For example, 
        vigorous fluvial action will separate elements that float easily from 
        those that do not (Voorhies 
        1969*1). By analyzing skeletal part frequencies, we can assess the 
        degree to which individual taxa have been affected by such factors. Although 
        it is seldom possible to determine the precise cause of incongruent element 
        frequencies, we may identify the most probable or predominant influences. 
      37 
        For this analysis, the remains for each taxon are assigned to one of seven 
        skeletal regions: cranial, axial, pectoral girdle, forelimbs (further 
        categorized into upper and lower), innominate, hind limbs (further categorized 
        into upper and lower), and phalanges (Table 
        9). It should be noted that some specimens could not be assigned to 
        a specific region, primarily those identified simply as "metapodial" or 
        "sesamoid." Several patterns are immediately apparent. Predictably, regions 
        with fewer skeletal components (for example, the shoulder and pelvis) 
        have consistently lower specimen frequencies. Otherwise, most taxa with 
        substantial numbers of identified specimens (that is, more than 100) are 
        well represented in all skeletal regions. The small rodents present an 
        exception to this pattern, as no phalanges were identified for these taxa. 
        This is not surprising: these small elements are unlikely to be consistently 
        recovered during excavation, and, because of morphological similarities 
        across different families, little effort was made to precisely identify 
        phalanges of small mammals during analysis. 
      38 
        The family Geomyidae (pocket gophers) displays a clearly skewed element 
        distribution: elements of the cranial region far outnumber those of all 
        other skeletal regions. This phenomenon has been documented at other sites 
        in the Southwest by researchers who have argued that it represents evidence 
        of human procurement of these animals (Shaffer 
        1992*1). Although this is certainly possible, it seems more probable 
        that the frequencies reflect recovery and identification biases. The only 
        elements in the skeleton of a pocket gopher that are likely to be collected 
        consistently are the cranium and, perhaps, the larger long bones (humerus, 
        tibia, and femur) and innominate, although even these can fall through 
        ΒΌ-inch mesh. Axial elements and phalanges, if recovered, are unlikely 
        to be identified precisely and, consequently, will most often be classified 
        as small mammal or small rodent. This phenomenon of higher-than-expected 
        frequencies of cranial elements is also evident to a lesser extent among 
        the slightly larger woodrat (Neotoma) and squirrel (Sciuridae) 
        remains, where cranial and hind-limb elements predominate.  
      39 
        Axial elements (ribs, sternum, and vertebrae) are prominent among the 
        medium-artiodactyl remains. This may suggest differential transport of 
        carcass portions to the site. However, all other skeletal regions are 
        also well represented, so the common occurrence of axial elements probably 
        reflects a high degree of fragmentation of these remains, rather than 
        a significant human behavioral pattern. 
      40 
        Overall, the element frequencies at Yellow Jacket Pueblo suggest that 
        complete animal carcasses were deposited at the site. The observed 
        variability among element frequencies can be accounted for by commonly 
        recognized recovery, identification, and quantification biases. No strong 
        patterns indicative of selective transportation or distribution of specific 
        butchery units are evident for any of the taxa. 
      Intrasite Variation
      41 
        The distribution of faunal remains at Yellow Jacket Pueblo is examined 
        here in an attempt to identify spatial and/or chronological patterning 
        within the site. The distributions of the identified taxa are presented 
        by architectural block in Table 
        10; the analysis presented here focuses on the three major taxa presumed 
        to be of cultural significance: turkey, lagomorphs, and artiodactyls. 
        Throughout these comparisons, the term "turkey" is used to refer to the 
        sum of all remains identified as either Meleagris gallopavo or 
        "large bird." Because of the relatively small sample sizes, detailed spatial 
        analyses of the less common taxa are not likely to reveal meaningful patterns. 
        Likewise, architectural blocks that yielded fewer than 50 specimens of 
        the three major taxa considered here are excluded from these analyses. 
      42 
        The absolute and relative frequencies of the three major taxa recovered 
        from each architectural block are presented in Table 
        11. Although considerable variability is apparent between the architectural 
        blocks, much of this can be attributed to the small samples recovered 
        from several areas (that is, Blocks 800, 1000, 1100, 2000, 2100, 2200, 
        2300, and 3300). When these blocks are excluded from consideration, general 
        consistency among the architectural blocks is apparent. Collectively, 
        turkey and lagomorph remains dominate the block assemblages, accounting 
        for more than 80 percent of the identified remains in all blocks. Most 
        of the block assemblages contain less than 11 percent artiodactyl remains, 
        with two exceptions: Block 2400 (14.4 percent) and Block 1200 (17.0 percent). 
        There is, however, considerable variation in the relative frequencies 
        of turkey and lagomorphs. Turkey bones typically make up 30 to 55 percent 
        of the major taxa, although Blocks 900 and 1200 stand out with values 
        of 23.7 percent and 19.3 percent, respectively. Lagomorph frequencies 
        range between 35.8 percent and 69.5 percent, with no clear clustering 
        of values. There is no obvious association between the relative frequencies 
        of these two species and the spatial organization of the blocks. No clusters 
        of blocks are apparent in which a particular taxon is either dominant 
        or underrepresented. 
      43 
        Some of this variability, however, may be explained by chronology. Differences 
        in the relative abundance of artiodactyl remains could be related to the 
        occupational history of the site. Those architectural blocks and associated 
        cultural deposits that are believed to have been first established during 
        the late Pueblo II period (see "Chronology") 
        consistently display relatively high frequencies of artiodactyl remains 
        (Figure 1). Specifically, 
        Blocks 100, 500, 700, 2400, and 3200 have artiodactyl frequencies ranging 
        between 6.5 percent and 14.4 percent of the major taxa. Artiodactyl remains 
        are consistently less common among those architectural blocks that are 
        believed to represent primarily mid- to late Pueblo III occupations. This 
        latter group includes Blocks 200, 300, 400, 600, 900, 1200, and 2500. 
        The one major exception to this pattern is Architectural Block 1200 (the 
        great tower complex), which has the greatest abundance of artiodactyl 
        remains in the tested areas of the site, despite being among the latest 
        occupied architectural units. That the great tower complex has a relatively 
        unusual faunal assemblage may attest to the specialized function of this 
        architectural unit. It is also notable that the bi-wall "tower kiva" within 
        this complex also displays a higher-than-average abundance of artiodactyl 
        remains, when compared with other mid- to late Pueblo III structures. 
        As is discussed below, similar patterns have been documented at Sand Canyon 
        Pueblo, another very large Pueblo III site in the region. 
      44 
        In contrast to the artiodactyl remains, the frequencies of lagomorph and 
        turkey bones do not display obvious spatial or chronological patterning. 
        The relative abundance of these taxa appears to vary sporadically among 
        blocks and through time and may simply reflect normal sampling variability 
        in conjunction with factors associated with taphonomy and quantification. 
      45 
        When compared across all tested blocks, the relative frequencies of the 
        three major taxa vary somewhat with context. Nonstructural contexts were 
        found to contain a relatively high abundance of turkey remains (42.1 percent) 
        when compared with contexts from structures (32.7 percent). Conversely, 
        lagomorph and artiodactyl remains are on average more abundant among structures: 
        artiodactyl remains represent 12.0 percent of the major taxa recovered 
        from structures and 6.8 percent among nonstructural contexts; lagomorphs 
        represent 55.3 percent among structure contexts and 51.0 percent among 
        nonstructural contexts. We have documented this pattern at other Pueblo 
        III sites in the region, most notably Sand Canyon Pueblo (Muir 
        1999*2) and Castle Rock Pueblo (Driver 
        2000*1), and believe it may relate to either storage/disposal practices 
        or natural taphonomic factors (see paragraph 50 ). 
      Intersite Variation
      Yellow Jacket Pueblo Compared With Other Pueblo III Sites in the Central 
        Mesa Verde Region
      46 
        The authors have analyzed the faunal assemblages of three other Pueblo 
        III sites in the regionSand Canyon Pueblo (Muir 
        1999*2), Castle Rock Pueblo (Driver 
        2000*1), and Woods Canyon Pueblo (Driver 
        2002*3)and the results of these analyses provide a basis for 
        intersite comparisons (Table 12). 
        These comparisons are presented in more detail elsewhere (Muir 
        and Driver 2002*2) and will only be briefly summarized and discussed 
        here. 
      47 
        Overall, the Pueblo III faunal assemblage from Yellow Jacket is most similar 
        to that from Sand Canyon Pueblo, both in terms of composition and distribution 
        of the remains. In particular, the relative frequency and distribution 
        of artiodactyl remains are remarkably similar at the two sites. While 
        Pueblo III sites in the central Mesa Verde region typically yield assemblages 
        dominated by turkey and lagomorphs, only Sand Canyon and Yellow Jacket 
        pueblos contain substantial quantities of artiodactyl remains (Muir 
        and Driver 2002*2). In both assemblages, artiodactyls make up more 
        than 8 percent of the major taxa. These remains are found clustered in 
        association with particular structures at each site. Much like the analyses 
        performed for Yellow Jacket Pueblo, spatial analyses of faunal remains 
        from Sand Canyon Pueblo (Muir 
        1999*2) reveal a marked concentration of artiodactyl remains among 
        deposits associated with towers and a more uniform distribution of major 
        taxa throughout the remainder of the site. Artiodactyl remains are also 
        relatively abundant among the D-shaped bi-wall structure at Sand Canyon 
        Pueblo (Muir and Driver 2002*2). 
        Lagomorph remains are distributed broadly throughout Sand Canyon Pueblo 
        and are generally more common in structures than in midden areas. Turkey 
        remains are also common in both midden and structure deposits throughout 
        most of the site, but are relatively scarce within towers and the bi-wall 
        structure. 
      48 
        The faunal assemblages from Woods Canyon and Castle Rock pueblos are also 
        similar to the assemblage from Yellow Jacket Pueblo, though there are 
        some notable differences. Artiodactyl remains are relatively scarce at 
        both sites; they represent only 3.19 percent of major taxa identified 
        at Castle Rock and less than 0.5 percent at Woods Canyon. Turkey remains 
        are predominant in the Woods Canyon assemblage, representing more than 
        73 percent of the major taxa, whereas lagomorphs represent only 26 percent. 
        Relative frequencies of these taxa from Castle Rock Pueblo are very similar 
        to those from Yellow Jacket, with lagomorphs most abundant (57 percent), 
        followed by turkey remains (39 percent). 
      49 
        Intrasite analysis of the Woods Canyon fauna (Driver 
        2002*3) revealed only minor variations in the spatial distribution 
        of turkey remains and relatively uniform distribution of the other taxa. 
        The rim complex at Woods Canyon Pueblo, which contains several towers 
        and a D-shaped structure, did not display marked concentrations of any 
        particular taxa but, rather, was characterized by relative frequencies 
        typical of the site as a whole. The only significant spatial patterning 
        evident among the remains from Woods Canyon was revealed in comparisons 
        between depositional contexts, where lagomorph remains were found to be 
        more commonly associated with structures than with midden or courtyard 
        areas. Similarly, at Castle Rock Pueblo the only significant source of 
        intrasite variation was seen in the lagomorph remains, which were again 
        found to be more common in deposits in structures (Driver 
        2000*1). 
      Discussion
      50 
        The broad distribution of both lagomorph and turkey remains throughout 
        all four sites and their predominance in midden assemblages suggest that 
        these animals were used widely and commonly throughout the villages. Both 
        turkey and lagomorphs are documented as having been primary sources of 
        meat for many historic Pueblo peoples (Gnabasik 
        1981*1), and the distribution of these taxa appears to be consistent 
        with that of common daily food refuse (or domestic household refuse). 
        However, lagomorph remains are more prevalent in structure deposits than 
        they are in nonstructural and midden areas. This is in contrast to turkey 
        remains, which are less abundant in structures. As we have discussed elsewhere 
        (Driver 2000*1; Muir 
        1999*2; Muir and Driver 
        2002*2), these patterns may reflect a wide range of factors, including 
        consequences of gender roles, value, status, spiritual associations, abandonment 
        ritual, or domesticity. We believe, however, that two factors stand out 
        as particularly probable explanations. Differences in the storage of these 
        animals would account for the observed patterns. Specifically, lagomorphs 
        are documented ethnographically as being stored whole (complete with bones) 
        within roomblocks, while no such practice appears to be documented for 
        turkeys (Gnabasik 1981*1). 
        This may explain the greater relative abundance of lagomorph remains within 
        structures. Alternatively, natural introduction of lagomorph remains to 
        the structure contexts may be responsible for their apparent abundance. 
        Cottontails may find the crevices and shelters provided by collapsed stone 
        masonry to be ideal locations for dens and burrows. However, our taphonomic 
        analyses of the assemblages at all sites consistently suggest that the 
        presence of lagomorphs is primarily the result of cultural activities. 
      51 
        The predominance of lagomorph and turkey remains at Yellow Jacket Pueblo 
        is typical of Pueblo III sites in the central Mesa Verde region. Driver 
        (2002*4) has demonstrated that the archaeological faunal record displays 
        a long-term regional trend toward an increased dependence on turkey and 
        a corresponding decrease in the exploitation of large game from Basketmaker 
        II through Pueblo III times. Given this trend, the relatively high abundance 
        of artiodactyl remains found in association with particular structures 
        at Yellow Jacket and Sand Canyon pueblos is intriguing in that it may 
        provide insight into the social and economic organization of these large 
        villages. 
      52 
        Muir (1999*2) has argued 
        that the distribution of artiodactyl remains at Sand Canyon Pueblo suggests 
        that remains found among towers and other associated structures are related 
        to communal hunting activities; further, certain architectural blocks 
        acted as hunting or war society houses (or offices), where the spoils 
        of communal hunts were processed and stored. Alternatively, bones of artiodactyls 
        might have been disposed of in particular locations or structures, possibly 
        to protect them from scavengers. Such "respectful" treatment of the remains 
        of game animals was widespread in North America (Beaglehole 
        1936*1; Luckert 1975*1; 
        Tanner 1979*1; Underhill 
        1946*1). 
      53 
        Lipe and Ortman (2000*1) 
        have suggested that multiwall structures functioned as residences for 
        families who had considerable ceremonial and political influence within 
        the community. The abundance of artiodactyl remains associated with these 
        structures may indicate that the residents of bi-wall structures had preferential 
        access to wild game. As discussed above, no such correlations between 
        structure type and faunal distribution were documented at Woods Canyon 
        and Castle Rock pueblos. It appears that activities that resulted in the 
        creation of such concentrations at the larger sites did not occur at these 
        smaller pueblos. This may indicate fundamental differences, corresponding 
        to community size, in the nature and organization of hunting activities 
        or in the distribution of the spoils of such hunts (Muir 
        and Driver 2002*2). 
      Conclusions
      54 
        The faunal assemblage from Yellow Jacket Pueblo is typical of faunal assemblages 
        from large late Pueblo II/Pueblo III sites in the central Mesa Verde region. 
        Turkey and lagomorphs are the predominant taxa represented in the assemblage, 
        whereas the remains of artiodactyls are less common and become increasingly 
        scarce through the Pueblo III period. This pattern has been documented 
        as part of a gradual, long-term (Basketmaker IIPueblo III) decrease 
        in dependence on large game and a corresponding increase in dependence 
        on domesticated species, as well as on wild animals that frequented garden 
        plots as "pests" (Driver 2002*4). 
        Skeletal representation for most taxa suggests that the bones are the 
        remains of complete animals that were brought to the site for butchery, 
        processing, and consumption. The presence of rodent remains recovered 
        from the site appears to be primarily the result of natural taphonomic 
        processes rather than cultural activities. 
      55 
        The spatial distribution of remains within Yellow Jacket Pueblo is similar 
        to that observed at Sand Canyon Pueblo. The concentration of artiodactyl 
        remains within and around specific structures at these sites suggests 
        that the procurement of large game was a communal activity. The great 
        tower complex at Yellow Jacket Pueblo might have been a facility in which 
        communal hunts were organized and/or communally acquired game was redistributed. 
        Alternatively, the complex might have been the residence or office of 
        political or spiritual leaders within the community. A larger sample of 
        faunal materials from the great tower complex and similar structures at 
        other sites in the region would be valuable for evaluating these possibilities. 
      Acknowledgments 
      This research was conducted with the cooperation 
        and assistance of Crow Canyon Archaeological Center researchers and staff. 
        We would like to thank all of the individuals who have been involved in 
        the Yellow Jacket project, particularly Kristin Kuckelman, Donna Glowacki, 
        Scott Ortman, and Mark Varien. Thanks are also owed to the Department 
        of Archaeology at Simon Fraser University, which provided facilities and 
        support. This research has been funded in part by grants to Dr. Jon Driver 
        from the Social Sciences and Humanities Research Council of Canada and 
        Simon Fraser University. 
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